MALAT1 and colon adenocarcinoma: Both variants were enriched in the nuclear/nuclear envelope (NE) fraction of all five cell types tested (Fig. 1J and Supplementary Fig. S3C–G), thus agreeing with previous data in colon adenocarcinoma cells [19], and resembling the strictly nuclear lncRNA MALAT1 [47], serving as a positive control (Supplementary Fig. S3D–G).