Additionally, several hub genes identified by WGCNA, including ACAT1 (Acetyl-CoA acetyltransferase 1), LIPC (Lipase C, hepatic type), GPLD1 (Glycosylphosphatidylinositol specific phospholipase D1), ACOX1 (Acyl-CoA oxidase 1), LPIN2 (Lipin 2), LPL (Lipoprotein lipase), ECI2 (Enoyl-CoA delta isomerase 2), and ACAA1 (Acetyl-CoA acyltransferase 1), have been verified in other species and identified as related to fatty liver. Here, GPLD1 is linked to fatty liver disease.