Immune pathways were also enriched: Role of Osteoclasts in Rheumatoid Arthritis (FDR = 5.88 × 10−3; MMP12, ADAM9, NFKBIA), T-cell receptor and Interleukin-1 signaling (both FDR = 8.83 × 10−3; NFKBIA, PSME3). Here, MMP12 is linked to rheumatoid arthritis.