WT1 and Hydrocephalus: Furthermore, the epigenetic modifier Smarcc1, which is linked to hydrocephalus (89), Foxn1, which is linked to neural tube defect and anencephaly (197) as well as Wt1, a marker of the radial glial (neural progenitor) cells (198) showed unique activity in the reconstructed dorsal neural tube after neural crest departure (Fig. 4E).