UMPS and colorectal adenocarcinoma: The ospC4 codes for Mxi-Spa TTSS effectors are controlled by virB, while fliR and fliY code for flagella cluster I. ompD is an outer membrane precursor involved in the adhesion and invasion of colonic epithelial cells as shown in human Caco-2, a colorectal adenocarcinoma cell line [42].