In the tumor cells at metastatic locations, we detected meta-program IV (MAP2, GRIK2), a hallmark of neuron development, and meta-program I, the mesenchymal-neuroblast-like (MES-Nbt) state, which was enriched for genes related to the EMT such as ZEB1, ZEB2, and TEAD1. Here, ZEB1 is linked to neoplasm.