By expressing the fly-FUCCI [36] using GMR85G01-GAL4 (Fig 2I), we showed that BBB glial cells actively progressed through the cell cycle during early larval stages and continued to proliferate post-CW in wild-type and tumor brains (Figs 2J, 2K and S4I–S4J′′′). Here, LGALS4 is linked to neoplasm.