These included pain traits like multisite chronic pain (MAML3, DCC, OLFM4, ANAPC4) and dorsalgia (CAMKV, MAML3, BCL11A, DCC, DCAF5, CNNM2, RABGAP1L, ZKSCAN2, ANAPC4, ARHGAP15, DRD2, MDGA2, CELF4), sleep traits like insomnia (CAMKV, DCC, OLFM4, CNNM2, MVK, NUDT12), and canonical psychiatric disorders like depression (DCC, NUDT12) and schizophrenia (CNNM2, STK31, PPP2R2B). Here, BCL11A is linked to major depressive disorder.