These included pain traits like multisite chronic pain (MAML3, DCC, OLFM4, ANAPC4) and dorsalgia (CAMKV, MAML3, BCL11A, DCC, DCAF5, CNNM2, RABGAP1L, ZKSCAN2, ANAPC4, ARHGAP15, DRD2, MDGA2, CELF4), sleep traits like insomnia (CAMKV, DCC, OLFM4, CNNM2, MVK, NUDT12), and canonical psychiatric disorders like depression (DCC, NUDT12) and schizophrenia (CNNM2, STK31, PPP2R2B). The gene discussed is ZKSCAN2; the disease is depressive disorder.