For 6 of these genes, a role in inflammatory bowel disease is supported by other criteria: GWAS associations with nearby SNPs (ANKRD55, LPP, PDLIM7), experimental perturbation (CCDC50, VCAM1), association with measured protein levels (IL6, VCAM1), or drug effects (IL6, VCAM1). Here, LPP is linked to inflammatory bowel disease.