Similarly, fixed SNPs were found in the EPHA4, EYS, HS6ST1, TOX genes, which have been associated with retinal nerve development (chickens; Fiore et al. 2019), retinal disease (humans; Yang et al. 2020), olfactory disease (humans; Tornberg et al. 2011), and near-sightedness (humans; Wang et al. 2022), respectively. Here, HS6ST1 is linked to Abnormal retinal morphology.