The O-linked glycosylation of mucins pathway was enriched with nine key proteins, including B2GNT7, C1GALT1, GALNT1, GALNT4, GALNT5, GALNT6, GALNT7, MUC13, and MUC5AC, each demonstrating significant upregulation in IPMN compared to ND (Figure 5B). Here, GALNT6 is linked to pancreatic intraductal papillary-mucinous neoplasm.