A relevant signal was on OCU8, at position 13.75–13.77 Mb, encompassing collagen type II alpha 1 chain (COL2A1), a gene linked to type II collagenopathies characterised by skeletal alterations and various forms of dwarfism in humans and cattle (Daetwyler et al., 2014; Deng et al., 2016). Here, COL2A1 is linked to Severe short stature.