MAPT and supranuclear palsy, progressive, 1: Neuropathologically, sequences of PSP-related tau pathology are stratified for accumulation of neuronal, astroglial, and oligodendroglial abnormal tau (Kovacs et al., 2020) into six steps for postmortem diagnosis: step 1 is characterized by intraneuronal tau inclusions in the globus pallidus, subthalamic nucleus, and substantia nigra; step 2 by accumulation of intraneuronal tau in the midbrain tegmentum, medulla oblongata, and pontine base, and astroglial tau pathology in the striatum.