For the hypertension and atrial fibrillation pairing, signals implicating shared causal variants were seen on chromosomes 4, 7, 11, and 17, in the genes FGF5, HOXA13, in the region spanning LSP1 to TNNT3 and that spanning CYTH1 to USP36 respectively. This evidence concerns the gene USP36 and atrial fibrillation.