From the between populations result, the candidate genes that were found in this study were previously associated with multiple diseases such as the tetraamelia syndrome in cattle (R-spondin 2 – RSPO2) [89], osteopetrosis in Red Angus (Solute carrier family 4 member 2 – SLC4A2) [90], liver cancer (MBL associated serine protease 2 – MASP2) [84], and mastitis (BoLA) [91]. Here, RSPO2 is linked to liver cancer.