Under the linkage peak of Chr6 QTL for atherosclerosis, Ath54, 41 protein-coding genes were polymorphic between BALB and LP strains, including Jazf1, Creb5, Tril, Cpvl, Chn2, Wipf3, Scrn1, Fkbp14, Plekha8, Mturn, Znrf2, Nod1, Ggct, Gars1, Crhr2, Inmt, Mindy4, Aqp1, Adcyap1r1, Neurod6, Itprid1, Ppp1r17, Pde1c, Lsm5, Avl9, Kbtbd2, Fkbp9, Nt5c3, Ppm1k, Herc6, Pyurf, Lancl2, Vopp1, Abcg2, Herc3, Fam13a, Gprin3, Snca, Mmrn1, Ccser1, and Grid2. Here, WIPF3 is linked to atherosclerosis.