Of them, Ctnnd2, Dap, Marchf6, Cmbl, Sdc2, Cpq, Stk3, Vps13b, Cox6c, Grhl2, Fzd6, Cthrc1, Lrp12, and Zfpm2 showed associations or had functions related to atherosclerosis or obesity. Here, LRP12 is linked to obesity due to melanocortin 4 receptor deficiency.