Further genome-wide association studies (GWAS) identified variants in the genes TCF4, SLC4A11, TCF8, KANK4, LAMC1, ATP1B1, and LOXHD1 to be associated with the late-onset form of FECD (Afshari et al. 2017; Baratz et al. 2010). Here, ZEB1 is linked to Fuchs endothelial corneal dystrophy.