Among the neoepitopes that were differentially expressed in HCC, nine (six shown) came from MUC12, “mucin 12 cell surface associated” protein, seven (six shown) came from HRNR, “hornerin” protein and seven (five shown) came from OR4C5, “olfactory receptor family 4 subfamily C member 5′ protein (Fig. 2c, Supplementary Table S5a). Here, HRNR is linked to hepatocellular carcinoma.