Four HLA alleles are significantly and positively correlated with HLA-C*03:02, including HLA-DRB*03:01 (r2=0.29; susceptible to MS and T1D, protective for IBD), HLA-DQB*02:01 (r2=0.28; susceptible to CD and T1D, protective for IBD), HLA-DQA1*05:01 (r2=0.29; susceptible to CD), and HLA-DRB*03 (r2=0.29; susceptible to T1D) (Figure 3; Table S2). Here, HLA-C is linked to inflammatory bowel disease.