GLUL and supranuclear palsy, progressive, 1: Some of the most upregulated genes were astrocyte marker genes, such as GFAP and GLUL [22,23], but also genes associated with protein phosphorylation regulation, such as microtubule-associated proteins MAP1A and MAP1B, but not MAPT, which was previously reported to be downregulated in PSP astrocytes [3,24,25,26,27].