PPARA and cancer: Consequently, PRAT adipocytes adopt beige/brown morphology features and exhibit overexpression of brown/beige adipocyte markers, such as UCP1, peroxisome proliferator-activated receptor ɣ (PPAR-ɣ), CCAAT/enhancer binding protein α (c/EBPα), and peroxisome proliferator-activated receptor gamma coactivator-1α (PGC1α), in cancer [20].