In addition, our TWAS pinpointed many tissue-specific sarcoidosis-associated genes and found expressions of 14 genes (ABT1, BTN2A2, C2orf74, CCDC88B, FAM213A, MDH2, METTL21B, PRSS16, RP3-473L9.4, TCF19, TSFM, UBASH3A, UQCC2, ZSCAN26) associated with sarcoidosis across all three target tissues. This evidence concerns the gene MDH2 and sarcoidosis.