Among the nine cell states associated with fibroblasts, five (ADAMDEC1+ fibroblast, BMP4+ fibroblast, intestinal smooth muscle, myofibroblast, PI16+ fibroblast) were identified as NAT-enriched, and three (WNT5A+ fibroblast, pericyte, desmoplastic fibroblast) were identified as tumor-enriched. Here, BMP4 is linked to neoplasm.