Furthermore, WD/FG increased serum total cholesterol, adipose tissue weight, liver triglyceride content, and steatosis score that were accompanied by significant upregulation of lipogenic genes like Acaca and Fasn. CEL significantly decreased all these lipid metabolism parameters and both genes in the set 1 experiment, which is in accordance with other animal studies (Hu et al., 2018; Zhang et al., 2016). This evidence concerns the gene FASN and steatosis.