A multi-progenitor meta-analysis identified 34 novel loci (P < 5e-8) and newly discovered candidate genes, including tumor necrosis factor alpha-induced protein 3 (TNFAIP3) interacting protein 2 (TNIP2) and TNF receptor superfamily member 11 A (TNFRSF11A), confirming the important roles of the immune system and joint tissues in the etiology of RA [13]. Here, TNIP2 is linked to rheumatoid arthritis.