Considering the known function of TOR3A, MPEG1, LHX8, TTN, ATP1A1, PCDH1, WHAMM and NGRN genes, the rarity of the associated candidate variants, and the outcome of the in silico effect prediction, the identified candidate variants in these genes were assumed to represent plausible candidate causal variants for bovine SS. Here, TTN is linked to synovial sarcoma.