ARAF and tetrasomy 12p: T. gaoligongensis contains 4 PKS genes: 1 NR-PKS, 2 PR-PKS, and 1 HR-PKS, characterized by the following domain structures: TgPKS1 (SAT-KS-AT-PT-ACP-ACP-TE), TgPKS2 (KS-AT-DH-KR-ACP-TE), TgPKS3 (KS-AT-KR-ACP-TE), and TgPKS4 (KS-AT-DH-MT-ER-KR-ACP-TE).