Li et al. (2016) found through linkage and association analysis that a QTL SCN3-11 resistant to cyst nematode (race 3) can explain 5.8% of the resistance variation in RIL populations derived from Zhonghuang 13 (susceptible) and Zhongpin 03-5373 (resistant). Aggregating the rhg1 site can significantly improve resistance. Through gene structure and expression analysis, it was further demonstrated that Glyma.11G234500 at SCN3-11 locus was a new disease resistance gene GmSNAP11 homologous to GmSNAP18 at rhg1-b locus (Lakhssassi et al., 2017). Here, PPIB is linked to cyst.