ZC3H12A and papilloma: On the other hand, our results showed that the upregulation of miR-92b-3p and miR-196a-5p in Mcpip1eKO papillomas is not a direct consequence of human MCPIP1 enzymatic activity, as the levels of these miRNAs were not increased in human keratinocytes expressing a shRNA targeting MCPIP1.