Specifically, CFHR1 was associated with AMD, WARS1 was associated with abnormal lens morphology and cataracts, LCT was associated with abnormal retinal morphology and decreased total retina thickness, and CTSH was associated with abnormal retina outer nuclear layer morphology, decreased total retina thickness, and abnormal retina inner nuclear layer morphology. Here, WARS1 is linked to age-related macular degeneration.