TMEM18 and obesity due to melanocortin 4 receptor deficiency: The TMEM18 (chr1: 8,940,691–9,011,524 bp) gene detected in this study has been reported to be associated with growth traits and obesity in rats (Rask-Andersen et al., 2012), cattle (Ma et al., 2012), and humans (Almén et al., 2010; Haupt et al., 2010).