Applying these 9982 models to GWAS data using S-PrediXcan, one lncRNA gene CTD-3080P12.3 (also known as TERLR1) was associated with overall breast cancer risk at the Bonferroni-corrected significance of P < 5.0 × 10−6 (0.05/9982). Here, TERLR1 is linked to breast cancer.