Restoration of HDAC complexes in the condensates severely dampens the activation of SS18-SSX downstream genes such as PAX6 and NKX3-2 (Fig. 3c) and almost abolishes the tumorigenicity of SS18-SSX tested by EdU cell proliferation assay (Fig. 3d, e), colony formation assay (Fig. 3f, g) and tumor-bearing mouse models (Fig. 3h, Supplementary Fig. 7a, b). Here, HDAC9 is linked to neoplasm.