ZEB2 and neoplasm: In agreement with these results, we found that in PDAC patient tumours the combined expression of WNT ligands (WNT7A, WNT7B, WNT4, WNT10A, WNT3, WNT10B, WNT11) has a positive correlation (R = 0.41, p value = 1.8e-08) with the combined expression of key EMT-inducing transcription factors SNAI1, SNAI2, ZEB1, ZEB2 and TWIST1 (Fig. 4i).