CCL4 and nasopharyngeal carcinoma: Lai et al. (2010) showed the LMP1-induced production of macrophage inflammatory protein-1 alpha (MIP-1α/CCL3) and macrophage inflammatory protein-1 beta (MIP-1β/CCL4) in EBV-infected NPC cell lines requiring both NF-κB and JNK signalling pathways [110].