Among all the 153 lncRNAs, only TUG1, STAG3L5P-PVRIG2P-PILRB, PVT1, LINC01089, HCP5, GUSBP11, and AL450992.2 were markedly up-regulated in HCC (Fig. 4B–C). Here, TUG1 is linked to hepatocellular carcinoma.