Given diagnostic advancements using molecular genetics (see the Molecular Genetics section), the updated definition of grade 3 (anaplastic) meningiomas now extends beyond traditional morphological subtypes and non-morphological criteria to include the presence of a telomerase reverse transcriptase (TERT) promotor and/or homozygous CDKN2A and/or CDKN2B deletions. Here, TERT is linked to meningioma.