DICER1 and tarsal-carpal coalition syndrome: When investigating the larger set of 107 inherited genes conferring risk across cancer types15, three additional candidate genes containing variants with higher prevalence in TCC cats were identified: DICER1, FAH, and COL7A1. Interestingly, the same heterozygous genotype for sibling cancer cats (Maliha, Padma, Pavarti, Gorton) was shared in both DICER1 and FAH. FAH and COL7A1 have not previously been associated with bladder cancer in human; however, mutations within these genes are connected to other cancers.