Furthermore, FUTs family expression at mRNA levels were determined by RT-qPCR in HBV-HCC and adjacent tissues (Fig. 1G), revealed an elevated level of FUT1, FUT2, FUT4, FUT5, FUT8, FUT10, PIGT, PIGM and B4GALT3, as well as a reduced level of FUT3, FUT6, FUT7, FUT9, FUT11, FUT12 and PIGV in HBV-HCC tissues. Here, PIGV is linked to hepatocellular carcinoma.