ACKR1 and Balkan nephropathy: This observation is partly explained by the finding that in populations of African and Yemenite Jewish ancestry, BEN is strongly associated with a polymorphism in the atypical chemokine receptor 1 (ACKR1/DARC), which encodes the Fy/Duffy antigen, a surface receptor utilized by P. vivax to invade RBCs [19].