Eight of these associations were novel (LMNA, SMAD6, HSPB9, TMEM95, KLF1, TET2, NME3, KDM5B) and 5 genes have previously been linked to arrhythmias (SCN5A, TTN, RPL3L, PKP2, PMVK). Here, TTN is linked to cardiac arrhythmia.