HLA-DRB5 and sarcoidosis: Interestingly, the rs1964995 SNP located between HLA-DRB9 and HLA-DRB5, a risk SNP only in AAs in our study, was protective in a Swedish cohort with non-Lofgren sarcoidosis GWAS [47], but increased the risk of rheumatoid arthritis in ADs, but not in EDs [48].