TIM4+CLEC4F+ KCs were always located outside and around the liver tumour nodules in an endogenous tumour model with spontaneous metastases (KPC mice) (Fig. 2a), as well as in the orthotopic graft model (Extended Data Fig. 2a), and in short-term models after intraportal injection of five different carcinoma and melanoma cell lines (Extended Data Fig. 2b–g). Here, CLEC4F is linked to neoplasm.