In contrast, C2GNT2—which is responsible for the core 2, core 4, and branched polylactosamine structures (I-branches)—were highly expressed in the malignant meningioma cell lines (HKBMM and IOMM-Lee) compared with the benign meningioma cell lines (SUT-MG12 and SUT-MG14). Here, GCNT3 is linked to meningioma.