These included the B3GALNT2 gene on chromosome 1, mutated in case of CH, the PLOD1 and the MUTYH genes on chromosome 2, mutated in case of WFFS and CA, respectively, the GBE1 gene on chromosome 26, mutated in case of GBED and the syntaxin 17(STX17) gene on chromosome 25, causative for grey color coat. Here, MUTYH is linked to cyclic hematopoiesis.