STX4 and Parkinson disease: Seven of these genes were novel for their association with PD, while twelve of them (i.e. ARHGAP27, FAM200B, TMEM175, CD38, ZSWIM7, GPNMB, STX4, KANSL1, ADORA2B, KAT8, MMRN1, PRSS36) were previously identified by Alvarado et al.21, validating our approach.