Analysis of PacBio long-read sequencing data from the ALT breast cancer cell line SK-BR-326 also revealed an enrichment of outward TFs in this line as compared to the non-ALT cell lines COLO289T, HCT116, KM12, SW620 and SW837, for which long-read sequencing data were also available27,28 (Supplementary Fig. 4B, C, Supplementary Fig. 6). Here, GPT is linked to breast carcinoma.