The rs187199523 locus has a binding site for the transcription factor ZKSCAN5 and is located on a related gene (RP11-563D10.1) intron, and RP11-563D10.1 is a long noncoding RNA (lncRNA), which might be associated with cholangiocarcinoma [35]. Here, ZKSCAN5 is linked to cholangiocarcinoma.