KLRC4 and Behcet disease: ERAP1 variants preferentially conferred risk for BD in HLA-B*51-positive individuals (p-value = 0.0009)ERAP1 p.Arg725Gln homozygosity was related with BD with an odds ratio of 3.78 [95% CI 1.94–7.35] in HLA-B*51-positive people and 1.48 [95% CI 0.78–2.80] in negative individualsNew susceptibility loci detected at CCR1, STAT4, and KLRC4 for BD