The role of primary cilia in radial migration is still debated, but previous studies have identified 30 ciliopathy-related genes that impact cerebral cortex development, with knock-down of 17 genes resulting in disturbance of distinct steps of radial migration, including a transient multipolar stage in the lower IZ, multipolar-to-bipolar transition in the upper IZ, and radial glia-guided locomotion in the CP (Guo et al., 2015). This evidence concerns the gene CP and ciliopathy.